Nutlet micromorphology and its taxonomic significance in Lamium sect. Amplexicaule (Lamiaceae)

Çiğdem CAN ÖZKER, Zeynep ATALAY, Ferhat CELEP


In Lamium, species-level identification is usually very difficult, particularly in closely related species. Here, we investigated nutlet micromorphological characteristics of five taxa belonging Lamium sect. Amplexicaule (L. amplexicaule, L. aleppicum, L. eriocephalum subsp. eriocephalum, L. eriocephalum subsp. glandulosidens, L. macrodon) and additional data taken from literature for L. persepolitanum by stereo-binocular microscopy and scanning electron microscopy to show their taxonomic value. The size of nutlets varies from 2.12 mm (L. amplexicaule) to 4.27 mm (L. eriocephalum subsp. glandulosidens) in length, 1.13 mm (L. amplexicaule) to 2.22 mm (L. eriocephalum subsp. glandulosidens) in width. The basic shape of nutlets is oblong, oblong-ovate, obovate and triangular. Five nutlet sculpturing patterns are present namely reticulate, penta-hegzagonal reticulate, penta-hegzagonal colliculate, verrucate, verrucate with reticulate pattern. The results shown that the variation of nutlet surface sculpturing pattern, nutlet shape, size, and macromorphological characters provide useful diagnostic properties at species level.


Lamium, Lamiaceae, nutlet, taxonomy, Scanning Electron Microscopy

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